search-in-menu: preference variable

Default value: 0

This variable controls what Epsilon does when you press "?" during completion and then continue typing a response. If zero, Epsilon moves from the pop-up list of responses back to the prompt area, and editing keys like <Left> navigate in the response. If nonzero, Epsilon moves in the pop-up menu of names to the first name that matches what you've typed, and stays in the pop-up window. (If it can't find a match, Epsilon moves back to the prompt as before.)

More info:

Completion & Defaults (General)

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search-delete-match Variable Reference search-in-region